<?xml version="1.0" encoding="UTF-8"?>
<!DOCTYPE article PUBLIC "-//NLM//DTD JATS (Z39.96) Journal Publishing DTD v1.3 20210610//EN" "JATS-journalpublishing1-3.dtd">
<article article-type="review-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="ru"><front><journal-meta><journal-id journal-id-type="publisher-id">virbul</journal-id><journal-title-group><journal-title xml:lang="ru">Бюллетень ВИР</journal-title><trans-title-group xml:lang="en"><trans-title>VIR Bulletin</trans-title></trans-title-group></journal-title-group><issn pub-type="epub">3034-0063</issn><publisher><publisher-name>N.I. Vavilov All-Russian Institute of Plant Genetic Resources (VIR)</publisher-name></publisher></journal-meta><article-meta><article-id custom-type="elpub" pub-id-type="custom">virbul-9</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>ОБЗОРНЫЕ СТАТЬИ</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>REVIEW ARTICLE</subject></subj-group></article-categories><title-group><article-title>Низкотемпературное хранение семян. Становление метода и его роль в сохранении генетических ресурсов растений</article-title><trans-title-group xml:lang="en"><trans-title>Low-temperature storage of seeds. Establishment of the method and its role in plant genetic resources conservation</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><contrib-id contrib-id-type="orcid">https://orcid.org/0000-0001-8419-1518</contrib-id><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Филипенко</surname><given-names>Г. И.</given-names></name><name name-style="western" xml:lang="en"><surname>Filipenko</surname><given-names>G. I.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Галина Ивановна Филипенко, кандидат сельскохозяйственных наук, ведущий научный сотрудник</p><p>190000 Россия, Санкт-Петербург, ул. Большая Морская, 42, 44</p></bio><bio xml:lang="en"><p>Galina I. Filipenko, Cand. Sci. (Agriculture), Leading Researcher</p><p>42, 44, Bolshaya Morskaya Street, St. Petersburg, 190000 Russia;</p></bio><email xlink:type="simple">g.filipenko@vir.nw.ru</email><xref ref-type="aff" rid="aff-1"/></contrib></contrib-group><aff-alternatives id="aff-1"><aff xml:lang="ru"><institution>Федеральный исследовательский центр Всероссийский институт генетических ресурсов растений имени Н.И. Вавилова</institution><country>Россия</country></aff><aff xml:lang="en"><institution>N.I. Vavilov All-Russian Institute of Plant Genetic Resources</institution><country>Russian Federation</country></aff></aff-alternatives><pub-date pub-type="collection"><year>2024</year></pub-date><pub-date pub-type="epub"><day>04</day><month>07</month><year>2026</year></pub-date><volume>0</volume><issue>243</issue><fpage>50</fpage><lpage>56</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Филипенко Г.И., 2026</copyright-statement><copyright-year>2026</copyright-year><copyright-holder xml:lang="ru">Филипенко Г.И.</copyright-holder><copyright-holder xml:lang="en">Filipenko G.I.</copyright-holder><license xml:lang="ru" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>Данная работа распространяется под лицензией Creative Commons Attribution 4.0.</license-p></license><license xml:lang="en" license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://www.vir-bulletin.ru/jour/article/view/9">https://www.vir-bulletin.ru/jour/article/view/9</self-uri><abstract><p>Около 90% генетических ресурсов культурных растений Земли сохраняется в виде коллекций семян. Низкотемпературное хранение образцов семян позволяет уменьшить число пересевов, поддерживать жизнеспособность семян на высоком уровне длительные сроки. В статье обсуждаются исследования влияния различных факторов на долговечность семян, ставшие методической основой при организации низкотемпературного хранения семян в генных банках растений. Рассмотрены случаи чрезвычайного долголетия семян, расширяющие наши представления о возможностях живых организмов. Приведены сведения из стандартов генных банков (Genebank Standards, 1994, Genebank Standards…, 2014) о выделении нескольких типов коллекций (активная, базовая, дублетная), что позволяет рационально организовать хранение и использовать имеющийся материал, а также о современных требованиях к условиям низкотемпературного хранения.</p></abstract><trans-abstract xml:lang="en"><p>About 90% of the global cultivated plant genetic resources are preserved as seed collections. The low-temperature storage of seed accessions allows for the reduction of the number of regenerations and for long term maintenance of high level seed viability. The article considers studies of the influence of various factors on seed longevity, which became the methodological basis for organizing low-temperature seed storage in plant genebanks. The presented cases of extreme seed longevity expand our knowledge of the capabilities of living organisms. The Genebank Standards (1994) and Genebank Standards for Plant Genetic Resources for Food and Agriculture (2014) provide information on the subdivision of collections into active, basic, and duplicate ones, which allows rational organization of storage and use of available material, as well as describe modern requirements to low-temperature storage conditions.</p></trans-abstract><kwd-group xml:lang="ru"><kwd>генетические ресурсы растений</kwd><kwd>стандарты генных банков</kwd><kwd>условия хранения</kwd><kwd>всхожесть семян</kwd><kwd>влажность семян</kwd></kwd-group><kwd-group xml:lang="en"><kwd>plant genetic resources</kwd><kwd>genebank standards</kwd><kwd>storage conditions</kwd><kwd>seed germination</kwd><kwd>seed moisture</kwd></kwd-group><funding-group><funding-statement xml:lang="ru">Работа выполнена в рамках реализации Программы развития Национального центра генетических ресурсов растений по соглашению с Минобрнауки России от 15 февраля 2024 года № 075-02-2024-1090.</funding-statement><funding-statement xml:lang="en">The work was carried out within the framework of the implementation of the Development Program of the National Center for Plant Genetic Resources under the Agreement with the Ministry of Education and Science of Russia dated February 15, 2024 No. 075-02-2024-1090.</funding-statement></funding-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Bailly C. Active oxygen species and antioxidants in seed biology. Seed Science Research. 2007;14 (2):93-107. DOI: 10.1079/SSR2004159</mixed-citation><mixed-citation xml:lang="en">Bailly C. Active oxygen species and antioxidants in seed biology. Seed Science Research. 2007;14 (2):93-107. DOI: 10.1079/SSR2004159</mixed-citation></citation-alternatives></ref><ref id="cit2"><label>2</label><citation-alternatives><mixed-citation xml:lang="ru">Becquerel P. La longevite des graines macrobiotiques. Comptes rendus de l'Académie des Sciences. 1934;194:2158-2159.</mixed-citation><mixed-citation xml:lang="en">Becquerel P. La longevite des graines macrobiotiques. Comptes rendus de l'Académie des Sciences. 1934;194:2158-2159.</mixed-citation></citation-alternatives></ref><ref id="cit3"><label>3</label><citation-alternatives><mixed-citation xml:lang="ru">Choudhury A., Bordolui S.K. Concept of seed deterioration: reason, factors, changes during deterioration and preventive measures to overcome seed degradation. American International Journal of Agricultural Studies. 2023;7(1):41-56. DOI: 10.46545/aijas.v7i1.291</mixed-citation><mixed-citation xml:lang="en">Choudhury A., Bordolui S.K. Concept of seed deterioration: reason, factors, changes during deterioration and preventive measures to overcome seed degradation. American International Journal of Agricultural Studies. 2023;7(1):41-56. DOI: 10.46545/aijas.v7i1.291</mixed-citation></citation-alternatives></ref><ref id="cit4"><label>4</label><citation-alternatives><mixed-citation xml:lang="ru">Corbineau F. Markers of seed quality: from present to future. Seed Science Research. 2012;22(1):61-68.</mixed-citation><mixed-citation xml:lang="en">Corbineau F. Markers of seed quality: from present to future. Seed Science Research. 2012;22(1):61-68.</mixed-citation></citation-alternatives></ref><ref id="cit5"><label>5</label><citation-alternatives><mixed-citation xml:lang="ru">Delouche J.C., Baskin C.C. Accelerated aging techniques for predicting relative storability of seed lots. Seed Science and Technology. 1973;1(2):427-452.</mixed-citation><mixed-citation xml:lang="en">Delouche J.C., Baskin C.C. Accelerated aging techniques for predicting relative storability of seed lots. Seed Science and Technology. 1973;1(2):427-452.</mixed-citation></citation-alternatives></ref><ref id="cit6"><label>6</label><citation-alternatives><mixed-citation xml:lang="ru">Ellis R.H., Hong T.D. Temperature sensitivity of the low-moisture-content limit to negative seed longevity-moisture content relationships in hermetic storage. Annals of Botany. 2006;97(5):785-791.</mixed-citation><mixed-citation xml:lang="en">Ellis R.H., Hong T.D. Temperature sensitivity of the low-moisture-content limit to negative seed longevity-moisture content relationships in hermetic storage. Annals of Botany. 2006;97(5):785-791.</mixed-citation></citation-alternatives></ref><ref id="cit7"><label>7</label><citation-alternatives><mixed-citation xml:lang="ru">Ewart A.J. On the longevity of seeds. Proceedings of the Royal Society of Victoria. 1908;21(1):1-210.</mixed-citation><mixed-citation xml:lang="en">Ewart A.J. On the longevity of seeds. Proceedings of the Royal Society of Victoria. 1908;21(1):1-210.</mixed-citation></citation-alternatives></ref><ref id="cit8"><label>8</label><citation-alternatives><mixed-citation xml:lang="ru">FAO. The Second Report on the State of the World’s Plant Genetic Resources for Food and Agriculture. Rome: Commission on genetic resources for food and agriculture FAO; 2010. URL: http://www.fao.org/4/i1500e/i1500e.pdf [accessed Dec. 01, 2024].</mixed-citation><mixed-citation xml:lang="en">FAO. The Second Report on the State of the World’s Plant Genetic Resources for Food and Agriculture. Rome: Commission on genetic resources for food and agriculture FAO; 2010. URL: http://www.fao.org/4/i1500e/i1500e.pdf [accessed Dec. 01, 2024].</mixed-citation></citation-alternatives></ref><ref id="cit9"><label>9</label><citation-alternatives><mixed-citation xml:lang="ru">Genebank Standards. 1994. Food and Agriculture Organization of the United Nations. Rome: International Plant Genetic Resources Institute; 1994. URL: https://cropgenebank.sgrp.cgiar.org/images/file/learning_space/genebank_standards.pdf [accessed Dec. 01, 2024].</mixed-citation><mixed-citation xml:lang="en">Genebank Standards. 1994. Food and Agriculture Organization of the United Nations. Rome: International Plant Genetic Resources Institute; 1994. URL: https://cropgenebank.sgrp.cgiar.org/images/file/learning_space/genebank_standards.pdf [accessed Dec. 01, 2024].</mixed-citation></citation-alternatives></ref><ref id="cit10"><label>10</label><citation-alternatives><mixed-citation xml:lang="ru">Genebank Standards for Plant Genetic Resources for Food and Agriculture. Rome: FAO; 2014. URL: https://www.fao.org/4/i3704e/i3704e.pdf [accessed Dec. 01, 2024].</mixed-citation><mixed-citation xml:lang="en">Genebank Standards for Plant Genetic Resources for Food and Agriculture. Rome: FAO; 2014. URL: https://www.fao.org/4/i3704e/i3704e.pdf [accessed Dec. 01, 2024].</mixed-citation></citation-alternatives></ref><ref id="cit11"><label>11</label><citation-alternatives><mixed-citation xml:lang="ru">Gomez-Campo C. Erosion of genetic resources within seedbanks: the role of seed containers. Seed Science Research. 2006;16:291-294.</mixed-citation><mixed-citation xml:lang="en">Gomez-Campo C. Erosion of genetic resources within seedbanks: the role of seed containers. Seed Science Research. 2006;16:291-294.</mixed-citation></citation-alternatives></ref><ref id="cit12"><label>12</label><citation-alternatives><mixed-citation xml:lang="ru">Hong T.D., Ellis R.H. A protocol to determine seed storage behaviour. Rome: International Plant Genetic Resources Institute; 1996.</mixed-citation><mixed-citation xml:lang="en">Hong T.D., Ellis R.H. A protocol to determine seed storage behaviour. Rome: International Plant Genetic Resources Institute; 1996.</mixed-citation></citation-alternatives></ref><ref id="cit13"><label>13</label><citation-alternatives><mixed-citation xml:lang="ru">Justice O.L., Bass L.N. Principles and practices of seed storage. Washington: U.S. Government Printing Office; 1978.</mixed-citation><mixed-citation xml:lang="en">Justice O.L., Bass L.N. Principles and practices of seed storage. Washington: U.S. Government Printing Office; 1978.</mixed-citation></citation-alternatives></ref><ref id="cit14"><label>14</label><citation-alternatives><mixed-citation xml:lang="ru">Хорошайлов Н.Г. Национальное хранилище семян мировых растительных ресурсов на Кубани. Бюллетень научно-исследовательского института растениеводства имени Н.И. Вавилова. 1978;77:3-9.</mixed-citation><mixed-citation xml:lang="en">Khoroshailov N.G. National repository of seeds of world plant resources in Kuban (Natsional'noye khranilishche semyan mirovykh rastitel'nykh resursov na Kubani). Research Bulletin of the N.I. Vavilov Institute of Plant Industry. 1978;77:3-9. [in Russian]</mixed-citation></citation-alternatives></ref><ref id="cit15"><label>15</label><citation-alternatives><mixed-citation xml:lang="ru">Libby W.F. Radiocarbon dates, II. Science. 1951;114:291-196.</mixed-citation><mixed-citation xml:lang="en">Libby W.F. Radiocarbon dates, II. Science. 1951;114:291-196.</mixed-citation></citation-alternatives></ref><ref id="cit16"><label>16</label><citation-alternatives><mixed-citation xml:lang="ru">McDonald M.B. Seed deterioration: physiology, repair and assessment. Seed Science and Technology. 1999;27(1):177-237.</mixed-citation><mixed-citation xml:lang="en">McDonald M.B. Seed deterioration: physiology, repair and assessment. Seed Science and Technology. 1999;27(1):177-237.</mixed-citation></citation-alternatives></ref><ref id="cit17"><label>17</label><citation-alternatives><mixed-citation xml:lang="ru">Ohga I. The germination of century old and recently harvested Indian lotus fruits, with special references to the effect of oxygen supply. American Journal of Botany. 1926;13:754-759.</mixed-citation><mixed-citation xml:lang="en">Ohga I. The germination of century old and recently harvested Indian lotus fruits, with special references to the effect of oxygen supply. American Journal of Botany. 1926;13:754-759.</mixed-citation></citation-alternatives></ref><ref id="cit18"><label>18</label><citation-alternatives><mixed-citation xml:lang="ru">Porsild A.E., Harrington C.R., Mulligan G. Lupinus arcticus Wats. grown from seeds of Pleistocene age. Science. 1967;158:113-114.</mixed-citation><mixed-citation xml:lang="en">Porsild A.E., Harrington C.R., Mulligan G. Lupinus arcticus Wats. grown from seeds of Pleistocene age. Science. 1967;158:113-114.</mixed-citation></citation-alternatives></ref><ref id="cit19"><label>19</label><citation-alternatives><mixed-citation xml:lang="ru">Roberts E.H. Viability of seeds. London: Chapman and Hall Ltd.; 1972. 448 p.</mixed-citation><mixed-citation xml:lang="en">Roberts E.H. Viability of seeds. London: Chapman and Hall Ltd.; 1972. 448 p.</mixed-citation></citation-alternatives></ref><ref id="cit20"><label>20</label><citation-alternatives><mixed-citation xml:lang="ru">Roberts E.H. Predicting the storage life of seeds. Seed Science and Technology. 1973;1:499-514.</mixed-citation><mixed-citation xml:lang="en">Roberts E.H. Predicting the storage life of seeds. Seed Science and Technology. 1973;1:499-514.</mixed-citation></citation-alternatives></ref><ref id="cit21"><label>21</label><citation-alternatives><mixed-citation xml:lang="ru">Roberts E.H., Ellis R.H. Water and seed survival. Annals of Botany. 1989;63:39-52. DOI: 10.1093/oxfordjournals.aob.a087727</mixed-citation><mixed-citation xml:lang="en">Roberts E.H., Ellis R.H. Water and seed survival. Annals of Botany. 1989;63:39-52. DOI: 10.1093/oxfordjournals.aob.a087727</mixed-citation></citation-alternatives></ref><ref id="cit22"><label>22</label><citation-alternatives><mixed-citation xml:lang="ru">Royal Botanic Gardens. Seed Collection. URL: https://www.kew.org/science/collections-and-resources/collections/seed-collection [accessed Dec. 01, 2024].</mixed-citation><mixed-citation xml:lang="en">Royal Botanic Gardens. Seed Collection. URL: https://www.kew.org/science/collections-and-resources/collections/seed-collection [accessed Dec. 01, 2024].</mixed-citation></citation-alternatives></ref><ref id="cit23"><label>23</label><citation-alternatives><mixed-citation xml:lang="ru">Sallon S., Solowey E., Cohen Y., Korchinsky R., Egli M, Woodhatch I., Simchoni O., Kislev M.E. Germination, Genetics, and Growth of an Ancient Date Seed. Science. 2008;320(5882):1464. DOI: 10.1126/science.1153600</mixed-citation><mixed-citation xml:lang="en">Sallon S., Solowey E., Cohen Y., Korchinsky R., Egli M, Woodhatch I., Simchoni O., Kislev M.E. Germination, Genetics, and Growth of an Ancient Date Seed. Science. 2008;320(5882):1464. DOI: 10.1126/science.1153600</mixed-citation></citation-alternatives></ref><ref id="cit24"><label>24</label><citation-alternatives><mixed-citation xml:lang="ru">Sallon S., Solowey E., Gostel M.R., Egli M., Flematti G.R. Bohman B., Schaeffer P., Adam P., Weeks A. Characterization and analysis of a Commiphora species germinated from an ancient seed suggests a possible connection to a species mentioned in the Bible. Communications Biology. 2024;7(1):1-13. DOI: 10.1038/s42003-024-06721-5</mixed-citation><mixed-citation xml:lang="en">Sallon S., Solowey E., Gostel M.R., Egli M., Flematti G.R. Bohman B., Schaeffer P., Adam P., Weeks A. Characterization and analysis of a Commiphora species germinated from an ancient seed suggests a possible connection to a species mentioned in the Bible. Communications Biology. 2024;7(1):1-13. DOI: 10.1038/s42003-024-06721-5</mixed-citation></citation-alternatives></ref><ref id="cit25"><label>25</label><citation-alternatives><mixed-citation xml:lang="ru">Сеферова И.В., Филипенко Г.И., Забегаева О.Н., Бойко А.П., Некрасов А.Ю., Новикова Л.Ю. Всхожесть семян сои после низкотемпературного хранения и ее зависимость от погодно-климатических условий в местах репродукции образцов. Биотехнология и селекция растений. 2024;7(3): 42-52. DOI: 10.30901/2658-6266-2024-3-o6</mixed-citation><mixed-citation xml:lang="en">Seferova I.V., Filipenko G.I., Zabegaeva O.N., Boyko A.P., Nekrasov A.Yu., Novikova L.Yu. Germination of soybean seeds after low-temperature storage and its dependence on weather and climate conditions in the places of reproduction of accessions. Plant Biotechnology and Breeding. 2024;7(3):42-52. [in Russian]. DOI: 10.30901/2658-6266-2024-3-o6</mixed-citation></citation-alternatives></ref><ref id="cit26"><label>26</label><citation-alternatives><mixed-citation xml:lang="ru">Смоликова Г.Н., Медведев С.С. Каротиноиды семян: синтез, разнообразие и функции. Физиология растений. 2015;62(1):1-13. DOI: 10.7868/S0015330315010133</mixed-citation><mixed-citation xml:lang="en">Smolikova G.N., Medvedev S.S. Seed carotenoids: synthesis, diversity, and functions. Russian Journal of Plant Physiology. 2015;62(1):1-13. [in Russian]. DOI: 10.7868/S0015330315010133</mixed-citation></citation-alternatives></ref><ref id="cit27"><label>27</label><citation-alternatives><mixed-citation xml:lang="ru">Строна И.Г. Общее семеноведение полевых культур. Москва: Колос; 1966.</mixed-citation><mixed-citation xml:lang="en">Strona I.G. General seed science of field crops. Moscow.: Kolos; 1966. 464 p. [in Russian]</mixed-citation></citation-alternatives></ref><ref id="cit28"><label>28</label><citation-alternatives><mixed-citation xml:lang="ru">Shvachko N.А., Khlestkina E.K. Molecular genetic bases of seed resistance to oxidative stress during storage. Vavilov Journal of Genetics and Breeding. 2020;24(5):451-458. DOI: 10.18699/VJ20.47-o</mixed-citation><mixed-citation xml:lang="en">Shvachko N.А., Khlestkina E.K. Molecular genetic bases of seed resistance to oxidative stress during storage. Vavilov Journal of Genetics and Breeding. 2020;24(5):451-458. DOI: 10.18699/VJ20.47-o</mixed-citation></citation-alternatives></ref><ref id="cit29"><label>29</label><citation-alternatives><mixed-citation xml:lang="ru">Svalbard Global Seed Vault. URL: https://www.seedvault.no/ [accessed Dec. 01, 2024].</mixed-citation><mixed-citation xml:lang="en">Svalbard Global Seed Vault. URL: https://www.seedvault.no/ [accessed Dec. 01, 2024].</mixed-citation></citation-alternatives></ref><ref id="cit30"><label>30</label><citation-alternatives><mixed-citation xml:lang="ru">The Convention on Biological Diversity. Article 2. Use of Terms. 1993. URL: https://www.cbd.int/convention/articles/default.shtml?a=cbd-02 [accessed Dec. 01, 2024].</mixed-citation><mixed-citation xml:lang="en">The Convention on Biological Diversity. Article 2. Use of Terms. 1993. URL: https://www.cbd.int/convention/articles/default.shtml?a=cbd-02 [accessed Dec. 01, 2024].</mixed-citation></citation-alternatives></ref><ref id="cit31"><label>31</label><citation-alternatives><mixed-citation xml:lang="ru">Walters C. Understanding the mechanisms and kinetics of seed aging. Seed Science Research. 1998;8:223-244.</mixed-citation><mixed-citation xml:lang="en">Walters C. Understanding the mechanisms and kinetics of seed aging. Seed Science Research. 1998;8:223-244.</mixed-citation></citation-alternatives></ref><ref id="cit32"><label>32</label><citation-alternatives><mixed-citation xml:lang="ru">Walters-Vertucci C., Crane J., Vance N.C. Physiological aspects of Taxus brevifolia seeds in relation to seed storage characteristics. Physiologia Plantarum. 1996;98(1):1-12.</mixed-citation><mixed-citation xml:lang="en">Walters-Vertucci C., Crane J., Vance N.C. Physiological aspects of Taxus brevifolia seeds in relation to seed storage characteristics. Physiologia Plantarum. 1996;98(1):1-12.</mixed-citation></citation-alternatives></ref><ref id="cit33"><label>33</label><citation-alternatives><mixed-citation xml:lang="ru">Yashina S., Gubin S., Maksimovich S., Yashina A, Gakhova E., Gilichinsky D. Regeneration of whole fertile plants from 30,000-y-old fruit tissue buried in Siberian permafrost. PNAS. 2012;109(10):4008-4013. DOI: 10.1073/pnas.1118386109</mixed-citation><mixed-citation xml:lang="en">Yashina S., Gubin S., Maksimovich S., Yashina A, Gakhova E., Gilichinsky D. Regeneration of whole fertile plants from 30,000-y-old fruit tissue buried in Siberian permafrost. PNAS. 2012;109(10):4008-4013. DOI: 10.1073/pnas.1118386109</mixed-citation></citation-alternatives></ref><ref id="cit34"><label>34</label><citation-alternatives><mixed-citation xml:lang="ru">Жукова Н.В., Хорошайлов Н.Г. Методические указания по длительному хранению семян / под ред. Н.Г. Хорошайлова. Ленинград: ВИР; 1981.</mixed-citation><mixed-citation xml:lang="en">Zhukova N.V., Khoroshailov N.G. Guidelines for long-term storage of seeds (Metodicheskiye ukazaniya po dlitel'nomu khraneniyu semyan). N.G. Khoroshailov (ed.). Leningrad: VIR; 1981. [in Russian]</mixed-citation></citation-alternatives></ref></ref-list><fn-group><fn fn-type="conflict"><p>The authors declare that there are no conflicts of interest present.</p></fn></fn-group></back></article>
